Proton pump

A proton pump is an mitochondrion, or other subcellular compartment.

Function

In cell respiration, the pumps grab protons from the matrix, the space between the two enclosing membranes of the organelle, and release the protons within the inner membrane. The confined protons create a difference or gradient in both electrochemical potential that acts as a kind of battery or reservoir of stored energy for the cell. The inner cell membrane functions in a similar way to a dam in a river. It blocks protons from drifting back into the matrix. Since the pumping action is against the gradient, it requires work (energy). The process is directly analogous to bicycling uphill or charging a battery (storing up potential energy). It is important to remember that the proton pump does not create energy. Instead, the gradient stores energy for the appropriate time.

Mobility

Some of the enzymes involved in the ubiquinone, travel along the matrix, like freight boats, navigating up and down the river.

Others such as NADH-Q reductase, act like ferryboats and cross the matrix. Enzymes that can cross the matrix may have a secondary role as proton pumps because they can deliver protons to the inner membrane.

Diversity

In humans

In ATPase proton pumps of other cellular membranes, ATP itself powers this transport.

The F_oF₁ proton channel temporarily opens in the inner membrane.

Non-humans

In bacteria and other ATP-producing organellesthan mitochondria, ATP-producing organelles, photosynthesis power the translocation of protons.

CF₁ ATP chloroplasts correspond to the human F_oF₁ ATP synthase in plants.

pigment used by archaea, most notably halobacteria.